Twentieth-century animal research psychologist and biologist. He is particularly remembered for his work on memory and learning processes in planarians, a variety of flatworms.
He taught in the University of Michigan's psychology department from 1963 until 1988.
He established both a scientific journal (the Journal of Biological Psychology) and a science-themed humor publication (the Worm Runner's Digest).
He lived most of his adult life in Ann Arbor, Michigan.
He was targeted by unabomber Ted Kaczynski and became partially deaf after opening an explosives-containing package from Kaczynski.