Evolutionary biologist and geneticist who worked to develop the mathematical basis of population genetics and evolutionary theory. He laid the foundations for the modern field of molecular evolution.
He was a student of Theodosius Dobzhanksy during his time studying at Columbia University.
He was the geneticist to first introduce the term "spandrel" into evolutionary theory.
His parents were Jewish.
He was a famous scientist and was born in New York; Herman Hollerith was also a well-known New York-born scientist.