Geneticist who contributed to the field of forensic science by developing advanced DNA profiling and fingerprinting methods.
He studied biochemistry at Merton College, Oxford and subsequently accepted a research fellowship at the University of Amsterdam.
He taught genetics courses at the University of Leicester.
He and his two siblings grew up in Oxford and Luton, England. He married Susan Miles in 1971.
He and fellow Englishman Paul Nurse both had acclaimed careers as geneticists.