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Reproductive scientist who discovered the soluble sperm factor and published the notable book Fertilization in Animals. He founded and edited Zygote, an academic journal that focuses on embryology.
He earned his Ph.D. in Cell Biology from the University of Manchester.
He is a Fellow both both the Society of Biology in London and the Royal College of Pathology.
He was born in Stockport, England.
He studied DNA, whose structure was first discovered by James D. Watson.